@....@ how does it work ?

Can somebody explain or point me to some documentation on what's going on with lines like :-
in my Makefile.in (built via glade & then automake) which then gets translated into :-
CFLAGS = -g -O2
by configure.
So how/where from does @CFLAGS@ become -g -O2 (medium-optimised) ?
My test program works OK, but I want to know what's going on in the configure. I have the Goat Book text but can't find the @...@ bit. I presume it's a substitution, but from where ?
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